The Better Angels of Our Nature: Why Violence Has Declined
the fruit and nuts that are unevenly distributed through the forest, they frequently split and coalesce into smaller groups ranging in size from one to fifteen. If one group encounters another group from a different community at the border between their territories, the interaction is always hostile. When the groups are evenly matched, they dispute the boundary in a noisy battle. The two sides bark, hoot, shake branches, throw objects, and charge at each other for half an hour or more, until one side, usually the smaller one, skulks away.
    These battles are examples of the aggressive displays that are common among animals. Once thought to be rituals that settle disputes without bloodshed for the good of the species, they are now understood as displays of strength and resolve that allow the weaker side to concede when the outcome of a fight is a foregone conclusion and going through with it would only risk injury to both. When two animals are evenly matched, the show of force may escalate to serious fighting, and one or both can get injured or killed. 12 Battles between groups of chimpanzees, however, do not escalate into serious fighting, and anthropologists once believed that the species was essentially peaceful.
    Jane Goodall, the primatologist who first observed chimpanzees in the wild for extended periods of time, eventually made a shocking discovery. 13 When a group of male chimpanzees encounters a smaller group or a solitary individual from another community, they don’t hoot and bristle, but take advantage of their numbers. If the stranger is a sexually receptive adolescent female, they may groom her and try to mate. If she is carrying an infant, they will often attack her and kill and eat the baby. And if they encounter a solitary male, or isolate one from a small group, they will go after him with murderous savagery. Two attackers will hold down the victim, and the others will beat him, bite off his toes and genitals, tear flesh from his body, twist his limbs, drink his blood, or rip out his trachea. In one community, the chimpanzees picked off every male in a neighboring one, an event that if it occurred among humans we would call genocide. Many of the attacks aren’t triggered by chance encounters but are the outcome of border patrols in which a group of males quietly seek out and target any solitary male they spot. Killings can also occur within a community. A gang of males may kill a rival, and a strong female, aided by a male or another female, may kill a weaker one’s offspring.
    When Goodall first wrote about these killings, other scientists wondered whether they might be freak outbursts, symptoms of pathology, or artifacts of the primatologists’ provisioning the chimps with food to make them easier to observe. Three decades later little doubt remains that lethal aggression is a part of chimpanzees’ normal behavioral repertoire. Primatologists have observed or inferred the killings of almost fifty individuals in attacks between communities, and more than twenty-five in attacks within them. The reports have come from at least nine communities, including ones that have never been provisioned. In some communities, more than a third of the males die from violence. 14
    Does chimpicide have a Darwinian rationale? The primatologist Richard Wrangham, a former student of Goodall’s, has tested various hypotheses with the extensive data that have been amassed on the demography and ecology of chimpanzees. 15 He was able to document one large Darwinian advantage and one smaller one. When chimpanzees eliminate rival males and their offspring, they expand their territory, either by moving into it immediately or by winning subsequent battles with the help of their enhanced numerical advantage. This allows them to monopolize access to the territory’s food for themselves, their offspring, and the females they mate with, which in turn results in a greater rate of births among the females. The community will also

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